Review of the Herrings of Iran (Family Clupeidae)

Brian W. Coad

Abstract

The systematics, morphology, distribution, biology, economic importance and conservation of the herrings (kilkas and shads) of Iran are described, the species are illustrated, and a bibliography on these fishes in Iran is provided. There are 9 native species in the genera Clupeonella, Alosa and Tenualosa in the Caspian Sea and rivers of southern Iran.

Keywords

Morphology, Biology, Alosa, Clupeonella, Tenualosa, Kilka, Shad.

Full Text:

PDF

References

Abbasi K. (2005). Comparative analysis of the diet of Alosa braschnikowi study in southeastern and southwestern coast of the Caspian Sea (Guilan, Mazandaran, Golestan). First Congress of Iran’s Biological Science, Tehran University, Karaj (abstract). (In Farsi)

Abbasi K., Moradkhah S., Sarpanah A.N. (2007). Identification and distribution of fish fauna in Siahdarvishan River (Anzali Wetland basin). Pajouhesh va Sazandegi, 19(1)(74): 27-39. (In Farsi)

Abbasi K., Sabkara J. (2004). Studying Alosa caspia caspia (Caspian shad) diet in southeastern Caspian Sea (Mazandaran and Golestan provinces). Iranian Journal of Biology, 17(3): 272-290. (In Farsi).

Abbasi K., Valipour A., Talebi Haghighi D., Sarpanah A., Nezami S. (1999). Atlas of Iranian Fishes. Gilan Inland Waters. Gilan Fisheries Research Centre, Rasht. 113 p. (In Farsi)

Abdoli A., Naderi M. (2009). Biodiversity of fishes of the southern basin of the Caspian Sea. Abzian Scientific Publications, Tehran. 243 p. (In Farsi)

Abdollapour Bereya H., Abbasi K., Sabkara J., Keyvan A. (2007). Studying Caspian shad (Alosa caspia caspia) diet in southwest coastal area of the Caspian Sea, Guilan Province waters. Iranian Scientific Fisheries Journal, 16(1): 115-128. (In Farsi)

Abdolmalaki S., Psuty I. (2007). The effects of stock enhancement of pikeperch (Sander lucioperca) in Iranian coastal waters of the Caspian Sea. ICES Journal of Marine Sciences, 64(5): 973-980.

Abtahi B., Taghavi Jolodar H., Yousefian M., Fazli H. (2004). Anatomical and histological study of ovary maturity stages of common kilka (Clupeonella delicatula). Pajouhesh va Sazandegi, 17(2): 47-54. (In Farsi)

Abtahi F., Taghavi H., Fazli H., Yousefian M. (2002). Fecundity and some biometrical characteristics of common kilka (Clupeonella delicatula) in Mazandaran Province. Iranian Journal of Marine Sciences and Technology, 1(2): 1-10. (In Farsi)

Afraei M.A. (2006). Identifcation and appointment distribution Clupeonella (genus Alosa) in the South Caspian Sea coastal. Iranian Fisheries Research Organization, Agriculture Research and Education Organization, Ministry of Jihad-e-Agriculture, Bandar Anzali. http://en.ifro.ir (abstract).

Afraei M.A., Parafkandeh Haghighi F., Janbaz A.A. (2006). Abundance and diversity of Clupeidae species in Mazandaran and Golestan coastal waters, north Iran. Iranian Scientific Fisheries Journal, 15(1): 21-32. (In Farsi)

Afraei Bandpei M.A. (2005). Age, growth, feeding regime and reproduction of Alosa braschnikowi in Iranian waters of the Caspian Sea (Mazandaran waters). M.Sc. Thesis, Islamic Azad University, North Tehran. 82p. (In Farsi)

Afraei Bandpei M.A., El-Sayed A-F.M., Pourgholam R., Nasrolahzadeh H., Valinassab T. (2012). Food and feeding habits of the Caspian marine shad, Alosa braschnikowi (Clupeidae) in the southern Caspian Sea. Cybium, 36(3): 411-416.

Afraei Bandpyi M.A., Salmani A., Khoshbavar Rostami H.R., Parafkandeh Hagbighy F. (2004). Biosystematic study of (Clupeidae) Alosa in Mazandaran and Golestan Province coastal waters. Iranian Journal of Fisheries Sciences, 13(3): 13-26. (In Farsi)

Agh N., Ehsani A., Jalili R., Ahmadian E. (2012). The effect of dietary blend vegetable oil on reproductive performance in rainbow trout (Oncorhynchus mykiss) broodstock. The First International Conference on Larviculture in Iran and International Workshop on Replacement of Fish/Meal Oil with Plant Sources in Aquatic Feed, Iran-Larvi 2012, 11-12 December 2012, Karaj, Iran. pp: 17-23.

Ahmadpour M., Ahmadpour M., Rezaei M.R., Karimzadeh G., Taheri K. (2012). Effect of the ctenophore invasion and adjust project of floating in the trend of change of kilka (Clupeidae) fishing in Iranian coasts of the Caspian Sea (1999-2009). Journal of Fisheries, 6(3): 55-60.

Akbari Miarkolae A., Khalesi M.K., Zoghi M., Rekabi S.M. (2014). Investigation on some biological and growth parameters of common kilka Clupeonella caspia Svetovidov, 1941 in southern Caspian Sea. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Akhondzadeh Basti A., Misaghi A., Zahraei Salehi T., Kamkar A. (2006). Bacterial pathogens in fresh, smoked and salted Iranian fish. Food Control, 17(3): 183-188.

Alam H. (No date). Fish and fisheries. In: E. Yarshater (Ed.). Encyclopædia Iranica. Bibliotheca Persica Press, New York. www.bibliothecapersica.com. downloaded 13 December 2006. 14 p

Alavi-Yeganeh M.S., Razavi S. (2016a). Morphological variation and taxonomic validation of gill raker characteristics in Caspian marine shad (Clupeidae: Alosa braschnikowi). Journal of Aquatic Ecology, 5(4): 122-127. (In Farsi)

Alavi-Yeganeh M.S., Razavi S. (2016b). Morphological characteristics of gill during development of Braschnikowi shad (Alosa braschnikowi: Clupeidae). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015. Pp: 92.

Alavi-Yeganeh M.S., Salehpour R., Tahmasebi M. (2013). Growth pattern and age distribution in three species of Caspian Sea herrings (Clupeidae: Alosa sp.) female along coast of Mazandaran Province. The First Iranian Conference of Ichthyology, Isfahan University of Technology, 15-16 May 2013, pp: 6.

Al-Baz A.F., Grove D.J. (1995). Population biology of sbour Tenualosa ilisha (Hamilton-Buchanan) in Kuwait. Asian Fisheries Science, 8: 239-254.

Al-Hassan L.A.J. (1982). The use of electrophoresis in the identification of fish stock, and its future application in the Arabian Gulf. Journal of the Faculty of Marine Science, Jeddah, 2: 81-84.

Al-Hassan L.A.J. (1988). Genetic variation and population structure in Nematalosa nasus and Thryssa hamiltonii from Iraqi waters. Bolletino del Museo Regionale di Scienze Naturali, Torino, 6(2): 639-646.

Al-Hassan L.A.J. (1993). Additional synopsis of biological data on Tenualosa ilisha. Marina Mesopotamica, supplement 2: 1-23 (In Arabic)

Al-Hassan L.A.J. (1999). Shad of the Shatt Al-Arab River in Iraq. The Shad Journal, Seattle, 4(2): 1-4.

Ali T.S., Mohamed A.R.M., Hussain N.A. (1998). The status of Iraqi marine fisheries during 1990-1994. Marina Mesopotamica, 13(1): 129-147.

Aliasghari M., Ghobadi Sh., Khodabakhsh E. (2011). Survey on population structure of bigeye kilka Clupeonella grimmi in Mazandarn Province coasts. Renewable Natural Resources Research, 2(3): 57-70.

Aliasghari M., Keramat Amirkolaie A., Parafkandeh Haghighi F. (2012). Investigation of some of the growth and reproductive parameters of common kilka Clupeonella caspia (Svetovidov, 1941) in southern Caspian Sea (Mazandaran zone). Journal of Fisheries (Iranian Journal of Natural Resources), 65(2): 205-217.

Aliasghari M., Parafkandeh Haghighi F. (2013). A comparative study on reproduction of kilka species (CLupeidae) in southeastern parts of the Caspian Sea. Iranian Journal of Fisheries Sciences, 12(3): 522-532.

Aliasghari M., Parafkandeh Haghighi F., Vatan Doust S., Ghasemnejhad H. (2011). Survey on growth parameters of anchovy kilka (Clupeonella engrauliformis) population in the southern Caspian Sea. Journal of Marine Biology, 2(4): 33-40. (In Farsi)

Al-Nasiri S.K., Al-Mukhtar M.A. (1988a). On the biology of sbour Hilsa ilisha (Hamilton) (Pisces, Clupeidae) from Ashar Canal, Basrah. Iraqi Journal of Agricultural Sciences "Zanco", 6(1): 97-106.

Al-Nasiri S.K., Al-Mukhtar M.A. (1988b). On the biology of sbour Hilsa ilisha (Hamilton) (Pisces, Glupeidae (sic)) from Ashar Canal, Basrah. Pakistan Journal of Zoology, 20(4): 321-328.

Al-Saad H.T. (1990). Hydrocarbon residues in fishes from Khor Al-Zubair, north west Arabian Gulf. Marina Mesopotamica, 5(1): 81-88.

Al-Saad H.T., Shamshoom S.M., Abaychi J.K. (1997). Hydrocarbons in fish from Shatt Al-Arab estuary and north-west Arabian Gulf following the Gulf oil spill of 1991. Marina Mesopotamica, 12(2): 213-231. (In Arabic)

Amodeo G. (1956). Report to the Government of Iraq on the possibilities of developing the sea fishing industry. Food and Agriculture Organization, Rome, (FAO/56/2/1327), Report No. 438: 20p.

Ashoori A., Naderi S., Barati A. (2012). Nestling diet of the grey heron, Ardea cinerea, in Siahkeshim, northern Iran (Aves: Ardeidae). Zoology in the Middle East, 57: 139-142.

Azizov A., Suleymanov S., Salavatian M. (2015). The features of the feeding of Caspian marineshad, Alosa braschnikowii (Borodin, 1904) in western part of the Caspian Sea. Caspian Journal of Environmental Sciences, 13(1): 77-83.

Badalov F.G. (1972). The food of Clupeonella delicatula (Nordm.) and C. engrauliformis (Borodin) in Bol'shoy Kyzylagach Bay in the Caspian. Journal of Ichthyology, 12(6): 1027-1030.

Bannai M.A.A., Muhammad E.T. (2016). Parasitic fauna of Tenualosa ilisha (Hamilton-Buchanan, 1822) as a biological indicator of fish immigraion in Iraqi waters. International Journal of Plant, Animal and Environmental Sciences, 6(3): 209-217.

Barati A. (2009). Diet and growth of chicks of the Great Cormorant, Phalacrocorax carbo, at Ramsar, northern Iran. Zoology in the Middle East, 46(1): 29-36.

Bartley D.M., Rana K. (1998). Stocking inland waters of the Islamic Republic of Iran. FAO Aquaculture Newsletter, 18: 16-19.

Barzegar M., Bozorgnia A., Youssefi M.R., Hosseinifard S.M. (2012). Determination of Alosa caspia persica parasites in fresh and brine water of Caspian Sea. World Journal of Fish and Marine Sciences, 4(2): 175-178.

Barzegar M., Raeisi M., Bozorgnia A., Jalali B. (2008). Parasites of eyes of fresh and brackish water fishes in Iran. Iranian Journal of Veterinary Research, Shiraz University, 9(3): 256-261.

Ben-Yami M. (1976). Fishing with Light. FAO Fishing Manuals. Fishing News Books, Farnham. 121p.

Berg L.S. (1948-1949). Freshwater fishes of the USSR and adjacent countries. Israel Program for Scientific Translations, Jerusalem (1962-1965). 3 volumes.

Bhaumik U. (2015). Review of global studies on food, growth and maturity profile of Indian shad (Tenualosa ilisha). International Journal of Current Research and Academic Review, 3(10): 127-139.

Bilio M., Niermann U. (2004). Is the comb jelly really to blame for it all? Mnemiopsis leidyi and the ecological concerns about the Caspian Sea. Marine Ecology Progress Series, 269: 173-183.

Blaber S.J.M., Milton D.A., Brewer D.T. (1997). Life history strategies of tropical estuarine shads genus Tenualosa (Clupeidae). Ninth International Congress of European Ichthyologists (CEI9) "Fish Biodiversity". Italy 1997 (Napoli-Trieste). Book of Abstracts, 14 p.

Blegvad H., Løppenthin B. (1944). Fishes of the Iranian Gulf. Einar Munksgaard, Copenhagen. 247p. (1999 translation into Farsi by E. Etemad and B. Mokayyer with supplement, Tehran University Publications No. 1744: 26 + 416p).

Borodin N. (1904). Novyi vid kaspiiskoi sel'di (Clupea engrauliformis n. sp.) [A new species of Caspian herring, Clupea engrauliformis n. sp.] Vestnik Rybopromyshlennosti, St. Petersburg, 19(6): 335.

Bulgakov G.P. (1926). K poznaniyu kaspiiskikh sel'dei. Opisanie novogo vida roda Caspialosa [To the knowledge of Caspian herrings. Description of a new species of the genus Caspialosa]. Byulleten' Sredneaziatskii Gosudarstvennyi Universitet, Taschkent, 13: 27-39.

Caddy J.F. (1984). Observations on prospects and constraints of development of Iranian fisheries. Food and Agriculture Organization, Rome, FI:TCP/IRA/2301 (Mf) Field Document 1: 30p.

Caspian Environmental Programme. (1998). Caspian Sea Environment National Report, Islamic Republic of Iran. Baku, Azerbaijan. Pagination various.

Caspian Environmental Programme. (2000). Biodiversity of the Caspian Sea. Baku, Azerbaijan. 122p.

Chang K.K. (1972). Composition of spawning stocks of the big-eyed shad [Alosa saposhnikovi (Grimm)] in the north Caspian. Journal of Ichthyology, 12(1): 46-54.

Coad B.W. (1997). Shad in Iranian waters. The Shad Journal, Seattle, 2(4): 4-8

Coad B.W. (1997). Status of shad in Iranian waters? The Shad Journal, Seattle, 2(2): 3.

Coad B.W., Hussain N.A., Ali T.S., Limburg K.E. (2003). Middle Eastern shads, p. 59-67. In: K.E. Limburg, J.R. Waldman (Eds.). Biodiversity, Status, and Conservation of the World's Shads. American Fisheries Society Symposium, 35: 369p.

Darvishi F. (2003). Study of feeding competition between invasion ctenophore (M. leidyi) and anchovy (C. engrauliformis) in the southern Caspian Sea (Mazandaran coast). Ph.D. Thesis, Islamic Azad University, Tehran. 179p. (In Farsi)

Darvishi F., Esmaili Sari A., Farshchi P. (2004). Dietary overlap between invasion Ctenophora (M. leidyi) and anchovy (C. engrauliformis) in the southern pars of Caspian Sea. International Journal of Environmental Science and Technology, 1(2): 89-96.

Das K., Al-Nasiri S.K., Shukri N.A., Sharma K.P. (1977). Biochemical studies on some commercially important fish of Shatt Al-Arab and the Gulf. In: Proceedings on the Handling, Processing and Marketing of Tropical Fish, London, July 1976. Tropical Products Institute, London. pp: 157-161.

Daskalov G.M., Mamedov E.V. (2007). Integrated fisheries assessment and possible causes for the collapse of anchovy kilka in the Caspian Sea. ICES Journal of Marine Science, 64(3): 503-511.

Dastan V., Abdi R., Movahedinia A.A., Salari Allabadi M.A. (2017). Study of gill and kidney tissue changes in Tenualosa ilisha during migration from sea to the Karun and Bahmanshir rivers. Iranian Scientific Fisheries Journal, 25(4): 53-61.

Dehbandi A., Motallebi A.A., Razavilar V., Pourgholam R. (2015). Effect of nicin z on some of spoilage chemical and bacterial properties in surimi of kilka (Clupeonella cultriventris caspia) stored in 4ºC. Iranian Scientific Fisheries Journal, 25(3): 41-56. (In Farsi)

Dmitriev N.A. (1947). Nekotorye dannye po biologii pekhleviiskogo (enzeliiskogo) puzanka (Caspialosa caspia knipowitschi Iljin) [Some biological data on the Pahlevi (Enzeli) shad (Caspialosa caspia knipowitschi Iljin)]. Zoologicheskii Zhurnal, 26(6): 559-560.

Dorvaj Z., Javadian S.R., Oveissipour M., Nemati M. (2013). Use of protein hydrolysates from Caspian Sea sprat (Clupeonella cultriventris) as a nitrogen source for bacteria growth media (Vibrio anguillarum, Bacillus licheniformis, Bacillus subtilis). Journal of Aquatic Animals and Fisheries, 4(15): 11-18.

Ebadi A.G., Shokrzadeh M. (2006). A survey and measurement of residues of lindane (organochlorine pesticides) in four species of the most consumed fish in the Caspian Sea (Iran). Toxicology and Industrial Health, 22(1): 53-58.

Ebrahimzadeh A., Nabawi, L. (1975). Survey of the parasites in the fishes of Karun River. Chamran University Publication, Ahvaz, 81: 1-35. (In Farsi)

Emadi, H. (1991). Biology and identification of Caspian kilka. Abzeeyan, Tehran, 2: 8-12. (In Farsi)

Ermolchev V.A., Ermolchev M.V., Besharat K. (1997). Results of the Russian-Iranian hydroacoustic survey of kilka stocks in the Southern Caspian to the south of the Astara-Gasan-Kuli line in 1995. PNIRO (Polar Research Institute of Marine Fisheries and Oceanography), Murmansk.

Ershad Langroudi H. (2004). Evaluation and comparison of durability of polycyclic aromatic hydrocarbon compounds (PAHs) and its relation to the lipid content in smoked fishes, silver carp (Hypophthalmichthys molitrix), Rutilus frisii kutum and herring (Alosa caspia). Ph.D. Thesis, Islamic Azad University, Tehran. 123p. (In Farsi)

Eschmeyer W.N., Ferraris C.J., Hoang M.D., Long D.J. (1996). A Catalog of the Species of Fishes. Preliminary version (Sept. 1996). www.calacademy.org/research/ichthyology/species.

Esmaeili H.R., Babai S., Gholamifard A., Pazira A., Gholamhosseini A., Coad B.W. (2015). Fishes of the Persis region of Iran: an updated checklist and ichthyogeography. Iranian Journal of Ichthyology, 2(3): 201-223.

Esmaeili H.R., Coad B.W., Mehraban H.R., Masoudi M., Khaefi R., Abbasi K., Mostafavi H., Vatandsoust S. (2014). An updated checklist of fishes of the Caspian Sea basin of Iran with a note on their zoogeography. Iranian Journal of Ichthyology, 1(3): 152-184.

Esmaeili H.R., Teimori A., Owfi F., Abbasi K., Coad B.W. (2014). Alien and invasive freshwater fish species in Iran: Diversity, environmental impacts and management. Iranian Journal of Ichthyology, 1(2): 61-72.

Esmaili Sari A., Farshchi P., Darvishi F. (2002). Feeding competition between invasive comb-jelly (M. leidyi) and anchovy kilka (C. engrauliformis) in south coast of the Caspian Sea. Iranian Journal of Marine Sciences, 1(4): 25-42. (In Farsi).

Faeed M., Mozafari N.A., Shojaee Arany A. (2006). Isolation and identification of bacteria and fungi spoilage in kilka meal production in Guilan Province. Iranian Journal of Fisheries Sciences, 14(4): 127-138. (In Farsi)

Fatemi M., Kaymaram F., Parafkandeh Haghighi F., Vosoughi G.H., Taghavi Motlagh S.A. (2009). Validation of back-calculation methods using otoliths to determine the length of anchovy kilka (Clupeonella engrauliformis). Iranian Journal of Fisheries Sciences, 8(2): 115-126.

Fazli H. (1990). Biology of genus Clupeonella in the Caspian Sea. National Conference on Fisheries Resources Management of the Caspian Sea, Babolsar. Iran Fisheries Organization, Sari. pp: 81-97. (In Farsi)

Fazli H. (2002). The study of catch and CPUE of kilkafishes in Mazandaran province (1996–2000). Virtual, 1(1). (In Farsi). (sic, abstract from Scientific Information Database, Tehran, http://en.journals .sid.ir/).

Fazli H. (2006). Monitoring (biology and catch) of kilka in Iranian commercial catch in 2001-2002. Iranian Fisheries Research Organization, Agriculture Research and Education Organization, Ministry of Jihad-e-Agriculture, Sari, http://en.ifro.ir (abstract).

Fazli H. (2007). Population dynamics and stock assessment of kilka (genus: Clupeonella) inIranian waters of the Caspian Sea. Ph.D. Thesis, Pukyong National University, Busan, South Korea. 145p.

Fazli H. (2011). Some environmental factors effects on species composition, catch and CPUE of kilkas in the Caspian Sea. International Journal of Natural Resources and Marine Sciences, 1(2): 157-164.

Fazli H., Asghar Janbaz A., Abdolmalaki Sh., Khedmati K. (2013). Kilka (Clupeonella) stocks status in Iranian waters of the Caspian Sea (1996-2011). Journal of Fisheries, 6(4): 33-42.

Fazli H., Besharat K. (1998). Kilka stock assessment using hydro-acoustic method and commercial catch monitoring, 1976-1977. Final report, Mazandaran Fisheries Research Center, Sari. 105p. (In Farsi)

Fazli H., Borani M.S., Janbaz A.A. (2005). Assessing the biological characteristics of Clupeonella grimi in Iranian commercial catch during 1997-2001 in the Caspian Sea. Iranian Journal of Fisheries Sciences, 13(4): 125-138. (In Farsi)

Fazli H., Borani M.S., Janbaz A.A. (2006). The biological characteristics of Clupeonella cultriventris caspia in southern coast and the impacts of Mnemiopsis leidyi invasion on the Caspian ecosystem. Pajouhesh va Sazandegi, 18(69): 87-96. (In Farsi)

Fazli H., Bourani M.S., Janbaz A.A., Kaymaram F., Amani G.A. (2003). Monitoring (biology and catch) of kilka in commercial catch in 2000. Iranian Fishery Research Organization, Tehran. 50p. (In Farsi).

Fazli H., Bourani M.S., Janbaz A.A., Roohi A. (2002). Kilka fishing and biological characteristics of anchovy before and after presence of Mnemiopsis leidyi in the Caspian Sea. First National Symposium on the Caspian Sea comb jelly (Mnemiopsis leidyi) 19-20 June 2002, Sari. (In Farsi)

Fazli H., Janbaz A.A., Bourani M.S., Kaymaram F., Amani G.A. (2004). Monitoring (biology and catch) of kilka in commercial catch in 2001-2002. Iranian Fishery Research Organization, Tehran. 60p. (In Farsi)

Fazli H., Janbaz A.A., Ghasemi S. (2017). Stock assessment and maximum sustainable yield of common kilka (Clupeonella cultriventris Borodin, 1904) in Iranian waters of the Caspian Sea. Iranian Scientific Fisheries Journal, 25(4): 87-99.

Fazli H., Janbaz A.A., Khedmati K., Parafkandeh F., Razavi Sayad B.A., Kor D., Taleshian H., Bagherzadeh F. (2006). Monitoring (biology and catch) of kilka in commercial catch in 2002-2005. Iranian Fishery Research Organization, Tehran. 50p. (In Farsi)

Fazli H., Janbaz A.A., Khedmati K., Taleshian H., Kor D., Bagherzadeh F., Bourani M. (2004). Kilka fishes monitoring (fishing and biology) in commercial fishing area (2002-2004). Iranian Fishery Research Organization, Tehran. 41p. (In Farsi)

Fazli H., Roohi A. (2002a). The impacts of Mnemiopsis leidyi on kilka resources in the southern Caspian Sea. Iranian Journal of Fisheries Sciences, 11(1): 63-72. (In Farsi)

Fazli H., Roohi A. (2002b). The impacts of Mnemiopsis leidyi on species composition, catch and CPUE of kilka in Iranian commercial catch. UNESCO, Caspian Floating University Research Bulletin, Astrakhan (KaspNIRKh), 3: 99-104. (In Russian)

Fazli H., Taghavi Jelodar H. (2013). Stocks status of kilka in Iranian waters of the Caspian Sea. International Journal of Environmental Resources Research, 1(2): 167-180.

Fazli H., Zhang C.I., Hay D.E., Lee C.W. (2009a). Multi-species approach for stock management of kilka fish (genus: Clupeonella) in Iranian waters. Iranian Journal of FisheriesSciences, 8(2): 141-154.

Fazli H., Zhang C.I., Hay D.E., Lee C.W. (2009b). Fishery biological characteristics and changes in the annual biomass of bigeye kilka (Clupeonella grimmi) in the Caspian Sea. Asian Fisheries Science, 22(3): 943-960.

Fazli H., Zhang C.I., Hay D.E., Lee C-W. (2009c). Stock assessment and management implications of anchovy kilka (Clupeonella engrauliformis) in Iranian waters of the Caspian Sea. Fisheries Research, 100(2): 103-108.

Fazli H., Zhang C.I., Hay D.E., Lee C.W., Janbaz A.A., Bourani M.S. (2007). Population dynamics and stock assessment of common kilka (Clupeonella cultriventris caspia) in Iranian waters of the Caspian Sea. Iranian Journal of Fisheries Sciences, 7(1): 47-70.

Fazli H., Zhang C-I., Hay D.E., Lee C-W., Janbaz A-A., Sayad Borani M. (2007). Population ecological parameters and biomass of anchovy kilka Clupeonella engrauliformis in the Caspian Sea. Fisheries Science, 73(2): 285-294.

Food and Agriculture Organization, Fisheries Department. (1996). Fisheries and aquaculture in the Near East and North Africa: situation and outlook in 1996. Food and Agriculture Organization, Rome, Fisheries Circular, 919: 37p.

Ghadirnejad H. (2003). Impact of the invasive ctenophore Mnemiopsis leidyi on the kilka fisheries Clupeonella engrauliformis in the southern Caspian Sea. Australian Society for Fish Biology Annual Conference, New Zealand (abstract) (www.asfb.org.au/pubs/2003/ 2003nz-51.htm, downloaded 12 January 2004).

Ghafleh Marammazi J., Eskandari G.R., Al-Mukhtar M.A., Kiabi B.H. (2004). Study of spawning season and spawning ground of soboor (Tenualosa ilisha, Ham. Bunch., (sic) 1822) during its migration in Khuzestan rivers. Iranian Journal of Fisheries Sciences, 4(1): 89-102, 124.

Ghayoumi R. (2009). Helminth parasites of Caspian kila (genus Clupeonella) as bioindicators of heavy metals. Ph.D. Thesis, Islamic Azad University, Tehran. 145p. (In Farsi)

Ghayoumi R., Malek M., Jamili S., Nabavi M., Motallebi A. (2009). Epizootiology of intestinal helminth parasites of Clupeonella spp. (Osteichthyes: Clupeidae) from the Caspian Sea, Iran. Bulletin of the European Association of Fish Pathologists, 29(4): 109-117.

Ghobeishavi A., Mousavi S.M., Yavari V., Kochanian P., Zakeri M. (2016). The innate immunity changes of the female anadromous hilsa shad, Tenulaosa ilisha, during spawning and post spawning season. Iranian Journal of Fisheries Sciences, 15(4): 1526-1539.

Ghotbi Jokanadan S.R., Alavi-Yeganeh M.S., Jamshidi S. (2014). Morphological comparison of two shad species, Alosa braschnikowi and Alosa saposchnikowii using morphometric and meristic characteristics in Guilan coasts. The Second Iranian Conference of Ichthyology, Faculty of Natural Resources, University of Tehran, Karaj, 7-8 May 2014 (abstract).

Ghotbi-Jokanadan S.R., Alavi-Yeganeh M.S., Jamshidi S. (2015). Length-weight and length-length relationships of four Alosa species along the southern Caspian Sea coast. Journal of Applied Ichthyology, 31(4): 814-815.

Harkonen T., Harding K.C., Wilson S., Baimukanov M., Dmitrieva L., Svensson C.J., Goodman S.J. (2012). Collapse of a marine mammal species driven by human impacts. PLOS One, 7(9): 1-9.

Hashemi H.S. (2001). Caspian Phoca (seal), Green Wave, Tehran, 5: 14-17. (In Farsi) (In English at www.netiran.com/Htdocs/WeeklyJournal/Social/wj00685.html, downloaded 5 November 2001).

Hashemi S., Eskandary G., Ansary H., Yooneszadeh M. (2011). Stock assessment and production of fish species in the Shadegan Wetland, Iran. World Journal of Fish and Marine Sciences, 3(6): 502-508.

Hashemi S., Mohammadi G., Eskandary G. (2010). Population dynamics and stock assessment of hilsa shad, (Tenualosa ilisha Hamilton-Buchanan, 1822) in coastal waters of Iran (northwest of Persian Gulf). Australian Journal of Basic and Applied Sciences, 4(12): 5780-5786.

Hashemi S.A., Ghorbani R., Kymaram F., Hossini S.A., Eskandari G., Hedayati A. (2014). Estimation of fish composition and catchability coefficient of gillnet in the Shadegan Wetland. Iranian Journal of Ichthyology, 1(1): 51-60.

Hashemi S.A., Ghorbani R., Kymaram F., Hossini S.A., Eskandari G., Hedayati A. (2015). Fish species composition, distribution and abundance in Shadegan Wetland. Fisheries and Aquaculture Journal, 6(2): 8p.

Hashemi S.A.R., Ansary H. (2012). Biomass and production of fish species in the Shadegan Wetland, Iran. Global Veterinaria, 9(2): 123-128.

Hashemi S.A.R., Mohammadi G., Safikhani H., Kashi M. (2009). Population dynamics and stock assessment of Hilsa shad and Tenualosa ilisha in Khuzestan Province coast (northwest of Gulf). The 6th Scientific Conference of Fisheries Resources, 3-4 March 2009, Basrah. pp: 121.

Hashemi S.A.R., Mohammadi G.H., Eskandari G.R. (2009). Growth and mortality parameters of hilsa shad, Tenualosa ilisha in Khuzestan Province coast. Journal of Marine Biology, 1(1): 64-77.

Hashemi Seyed A.R., Mohammadi G.H., Eskandari Gh.R., Kashi M.T. (2010). Population dynamics and stock assessment of hilsa shad, Tenualosa ilisha in Khuzestan Province coast. Journal of Fisheries, 3(4): 21-28. (In Farsi)

Heidari A., Paknejad S., Mousavi-Sabet H. (2015a). Morphometric and meristic characteristics of shad fish population (Alosa braschnikowii: Clupidae (sic)) from the Miyankaleh Gulf in the southeastern Caspian Sea. The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015. Pp: 23.

Heidari A., Paknejad S., Mousavi-Sabet H. (2015b). Length-weight relationships of shad fish (Alosa kessleri: Clupeidae), in Tonekabon fishing area, south of the Caspian Sea. The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015, pp: 24.

Hindi M.J., Sarhan H.R., Al-Shatty S.M.H. (1996a). Quality criteria of fresh carp, Cyprinus carpio and sbour, Tenualosa ilisha. 1 - The chemical composition. Marina Mesopotamica, 11(2): 251-261.

Hindi M.J., Sarhan H.R., Al-Shatty S.M.H. (1996b). Quality criteria of fresh carp, Cyprinus carpio and sbour, Tenualosa ilisha. 2 - Chemical indices. Marina Mesopotamica, 11(2): 263-272.

Hoestlandt H. (1991). The Freshwater Fishes of Europe. Volume 2. Clupeidae, Anguillidae. AULA-Verlag, Wiesbaden. 448p.

Holčík J., Oláh J. (1992). Fish, fisheries and water quality in Anzali Lagoon and its watershed. Report prepared for the project - Anzali Lagoon productivity and fish stock investigations. Food and Agriculture Organization, Rome, FI:UNDP/IRA/88/001 Field Document 2: 109p.

Hosseini S.S., Jeiran A., Aghili K., Rezaei Shirazi A. (2011). The survey morphometric and meristic of (Clupeidae) Alosa in Gilan Province coastal waters. Journal of Aquatic Animals and Fisheries, 2(7): 9-17.

Hussain N.A. (1997). Hilsa shad invade estuary in the Arabian Gulf. The Shad Journal, Seattle, 2(4): 2.

Hussain N.A., Ali T.S., Coad B. (2001). Middle Eastern shads. Paper presented at Shad 2001: A Conference on the Status and Conservation of Shads Worldwide, 20-23 May, 2001, National Aquarium, Baltimore, Maryland (abstract).

Hussain N.A., Jabir M.K., Yousif U.H. (1994). On the biology of sbour Tenualosa ilisha (Hamilton) in the Shatt Al-Arab River, south of Iraq, with notes on their distribution in the Arabian Gulf. Marina Mesopotamica, 9(1): 115-139.

Hussain N.A., Mohamed A-R.M., Al Noor S.A., Coad B., Mutlak F.M., Ibrahem M., Al-Sudani I.M., Mojer A.M., Toman A.J., Abdad M.A. (2006). Species composition, ecological indices, length frequencies and food habits of fish assemblages of the restored southern Iraqi marshes. Report by the University of Basrah, Basrah, Iraq. 114p.

Hussain S.A., Al-Mukhtar M.A., Al-Daham N.K. (1991). Preliminary investigation on fishery and some biological aspects of sbour, Hilsa ilisha, from Shatt al-Arab River, Iraq. Basrah Journal of Agricultural Science, 4(1, 2): 141-151.

Islam B.N., Talbot G.B. (1968). Fluvial migration, spawning, and fecundity of Indus River hilsa, Hilsa ilisha. Transactions of the American Fisheries Society, 97(4): 350-355.

Ivanov V.P., Katunin D.N. (2001). Report on the results of the Caspian Marine expedition. Caspian Regional Thematic Center for Management of Bioresources (CRTC MB), Caspian Environmental Programme, Baku, Azerbaijan. 2 volumes.

Jaafari O., Nasrolah-Pourmoghadam M., Shabani A. (2013). Comparison of some morphological features of Caspian marine shad (Clupeidae: Alosa braschnicowi (sic)) between regions of Sari and Anzali. The First Iranian Conference of Ichthyology, Isfahan University of Technology, 15-16 May 2013, pp: 29.

Jafari O., Shabani A., Kolangi Miandare H. (2014). Population structure of the Caspian shad (Alosa braschnikowi Borodin, 1904) in the southern coast of the Csspian Sea between Gomishan and Mainkaleh regions. Journal of Taxonomy and Biosystematics, 6(19): 15-26. (In Farsi)

Jabir M.K., Faris A.A. (1989). Fecundity of sbour, Tenualosa ilisha (Hamilton-Buchanan, 1822) in the Shatt al-Arab River, Basrah. Marina Mesopotamica, 4(2): 281-296.

Jafri S.I.H. (1994). The anadromous shad, Tenualosa ilisha: an overview of its fishery in Pakistan. Proceedings of First Symposium on Fish and Fisheries of Pakistan held at Department of Zoology, Government College, Lahore, Pakistan on October 19-20, 1991, pp: 33-42.

Jalili R., Agh N., Asgari R., Bagherzadeh Lakani F. (2012). Effects of replacing dietary fish meal with different protein sources on growth performance of rainbow trout (Oncorhynchus mykiss) fry. The First International Conference on Larviculture in Iran and International Workshop on Replacement of Fish/Meal Oil with Plant Sources in Aquatic Feed, Iran-Larvi 2012, 11-12 December 2012, Karaj, Iran, pp: 282-291.

Janbaz A.A., Abdolmalaki Sh. (2009). Age, growth and mortality of Clupeonella cultriventris caspia in coastal Mazandaran. Pajouhesh va Sazandegi, 21(4): 78-80. (In Farsi)

Janbaz A.A., Fazli H., Pourgholam R., Afraei Bandpei N. A., Nasrollahzadeh Saravi H., Abd Almaleki Sh. (2012). Reproduction, sexual maturity and fecundity of anchovy kilka (Clupeonella engrauliformis Svetovidov 1941) in Iranian waters of the Caspian Sea. Iranian Scientific Fisheries Journal, 20(4): 21-32. (In Farsi)

Janbaz A.A., Fazli H., Pourgholam R., Kaymaram F., Afraei Bandpei M.A., Abdolmaleki S., Khedmati K. (2012). Fishery and biological aspects of anchovy kilka (Clupeonella engrauliformis) in the southern Caspian Sea. Iranian Journal of Fisheries Sciences, 11(4): 796-806.

Janbaz A.A., Fazli H., Pourgholam R., Kaymaram F., Khedmati K., Parafkandeh F., Afraei Bandpei M.A. (2016). Age and growth of bigeye kilka (Clupeonella grimmi Kessler, 1877) in Iranian waters of the Caspian Sea. Iranian Journal of Fisheries Sciences, 15(4): 1410-1424.

Janbaz A.A., Kor D., Moghim M., Afraei M.A., Abdolmaleki S., Daryanabard G., Bagheri S., Khedmati K., Shabani K., Nahrevar M.R., Rastin R. (2010). Biological aspect of kilka (age, growth, feeding and reproduction) in southern Caspian Sea. Caspian Sea Ecology Research Center, Sari. 122p.

Jawad L.A., Al-Mukhtar M.A., Ahmed H.K. (2004). The relationship between haematocrit and some biological parameters of the Indian shad, Tenualosa ilisha (Family Clupeidae). Animal Biodiversity and Conservation, 27(2): 47-52.

Jolodar M.N., Abdoli A. (2004). Fish Species Atlas of South Caspian Sea Basin (Iranian Waters). Iranian Fisheries Research Organization, Teheran.110p. (In Farsi and English)

Jorfi E., Amini F., Ghoreyshi S.A., Mortezaei S.R.S. (2008). A study on population genetics of hilsa shad, Tenualosa ilisha, in Khoozestan, Iran using molecular method (RAPD). Iranian Scientific Fisheries Journal, 17(1): 31-44. (In Farsi)

Jorfi E., Farhad A., Reza M.S. (2009). A comparison between sobour (Tenualosa ilisha) of Iran and Iraq by molecular method. The 6th Scientific Conference of Fisheries Resources, 3-4 March 2009, Basrah, pp: 155.

Jorjani S., Khanipour A.A., Ghelichi A. (2014). Chemical composition and fatty acid profile of common kilka, Clupeonella cultriventris caspia. Caspian Journal of Environmental Sciences, 12(1): 119-128.

Jouladeh-Roudbar A., Vatandoust S., Eagderi S., Jafari-Kenari S., Mousavi-Sabet H. (2015). Freshwater fishes of Iran; an updated checklist. AACL Bioflux, 8(6): 855-909.

Kanazawa R.H. (1955). Fishes of Iraq and Iran. American Documentation Institute, ADIM 4612: 43-46.

Karimi N., Bani A., Abdolmalaki S. (2013). Reproductive condition and ovarian atresia in the common kilka (Clupeonella cultriventris) from the Caspian Sea years after a dramatic outbreak of the comb jelly fish (Mnemiopsis leidyi). Scientia Marina, 77(3): 501-509.

Karimi Iraj Z., Pourkhabbaz A.R., Hassanpour M., SinkaKarimi M.H. (2014). Bioaccumulation of heavy metals in tissues of Clupeonella cultriventris and Alosa caspia and their consumption risk assessment in the southern coast of Caspian Sea. Journal of Mazandaran University of Medical Sciences, 24(118): 99-110. (In Farsi)

Karimzadeh G. (2011a). Study of natural and fishing mortality and exploitation rate of common kilka Clupeonella cultriventris in southeast part of the Caspian Sea (Babolsar). Annual Review and Research in Biology, 1(3): 57-67.

Karimzadeh G. (2011b). Study of the natural and fishing mortality and exploitation rates of bigeye kilka (Clupeonella grimmi) in the southeast part of the Caspian Sea (Babolsar). African Journal of Agricultural Research, 6(3): 676-680.

Karimzadeh G., Gabrielyan B., Fazli H. (2010). Population dynamics and biological characteristics of kilka species (Pisces: Clupeidae) in the southeast coast of the Caspian Sea. Iranian Journal of Fisheries Sciences, 9(3): 422-433.

Keivany Y., Nasrollahzadeh G.A. (1990). A report on fishing and fishery industries in Iran. Gorgan University College, Gorgan, Iran. MS, 22 p. (In Farsi, English abstract)

Khanedan N., Motalebi A.A., Khanipour A.A., Koochekian Sabour A., Seifzadeh M., Hasanzati Rostami A. (2011). Effects of different concentrations of sodium alginate as an edible film on chemical changes of dressed kilka during frozen storage. Iranian Journal of Fisheries Sciences, 10(4): 654-662.

Khanipour A.A., Fathi S., Fahim Dejban Y., Zareh Gashti G. (2013). Chemical indicators for spoilage and shelf-life of the consolidated burgers (kilka-silver carp) during cold storage at -18°C. Iranian Scientific Fisheries Journal, 22(3): 41-48.

Khanipour A.A., Jorjani S., Soltani M. (2014). Chemical, sensory and microbial quality changes of breaded kilka (Clupeonella cultriventris) with tempura batter in production stage and during frozen storage. International Food Research Journal, 21(6): 2421-2430.

Khedmati Bazkiaei K., Khara H., Abdolmaleki S., Vahabzadeh H. (2013). Some of biology characteristic and reproduction of Clupeonella culriventris in the coastal waters of Anzali Port. The First Iranian Conference of Ichthyology, Isfahan University of Technology, 15-16 May 2013, pp: 39.

Khedmati Bazkiaei K., Khara H., Abdolmaleki S., Vahabzadeh H. (2014). Study on feeding of common kilka (Clupeonella caspia) in the Caspian Sea, Anzali shore. Journal of Marine Biology, 6(21): 13-22. (In Farsi)

Khorashadizadeh M.A., Abtahi B., Kazemi R., Fazli H. (2006). Anatomical and histological study of ovary development stages of big-eye kilka (Clupeonella grimmi) in Babolsar area, Iran. Iranian Journal of Fisheries Sciences, 15(3): 61-74. (In Farsi)

Khoshkhoo J. (2008). Study on the sustainability of FPC produced from kilka (Clupeonella engrauliformis, C. grimmi and C. cultriventris) in two vacuum packaging and modified atmosphere packaging at different environmental factors. Ph.D. Thesis, Islamic Azad University, Tehran. 193p. (In Farsi)

Khoshkhoo Zh., Motalebi A., Khanipour A.A., Karimi Firozjaee H., Nazemi M., Mahdabi M. (2010). Study on changes of protein and lipid of fish protein concentrate (FPC) produced from kilkas in VP and MAP packages at light and darkness conditions durng six months. International Journal of Environmental Science and Development, 1(1): 101-106.

Khoshkhoo Zh., Motalebi A.A., Razavilar V., Khanipour A.A. (2012). Protein and lipid changes of FPC produced from Caspian Sea kilkas in VP and MAP during storage at different temperatures. Iranian Journal of Fisheries Sciences, 11(2): 338-346.

Kiabi B.H., Abdoli A., Naderi M. (1999). Status of the fish fauna in the South Caspian Basin of Iran. Zoology in the Middle East, 18: 57-65.

Kideys A.E., Roohi A., Bagheri S., Finenko G., Kamburska L. (2005). Impacts of invasive ctenophores on the fisheries of the Black Sea and Caspian Sea. Oceanography, 18(2): 76-85.

Kinneir J.M. (1813). A geographical memoir of the Persian Empire, accompanied by a map. John Murray, London. 486p.

Knipovich N.M. (1921). Hydrological investigations in the Caspian Sea in the years 1914-1915. In: N.M. Knipovich (ed.). Reports of the Caspian Expedition in the years 1914-1915, Gosudarstvennoe Izdatelstvo, Petersburg. 943p. (In Russian)

Kohani M., Moradinasab Gh., Daliri M., Paighambari S. Y., Bibak M. (2012). Temporal variation of fishing and stock status of kilka fish (Clupeonella) in the fishing grounds of Guilan province during the years 1989-2009. The First National Conference on Catch and Exploitation of Aquatic Animals Stocks, Islamic Azad University of Abadan, December 2012, pp: 99-106. (In Farsi)

Koochekian Sabour A., Moini S. (2009). Producing fish sauce from Caspian kilka. Iranian Journal of Fisheries Sciences, 8(2): 155-162.

Kosarev A.N., Yablonskaya E.A. (1994). The Caspian Sea. SPB Academic Publishing, The Hague. 259p.

Kottelat M., Freyhof J. (2007). Handbook of European freshwater fishes. Kottelat, Cornol, Switzerland and Freyhof, Berlin, Germany. 646p.

Krylov V.I. (1984). An estimate of the effect of the Caspian seal (Pusa caspica) on fish populations. Canadian Translation of Fisheries and Aquatic Sciences, 5066: 15p.

Kushnarenko A.L. (1986). Ecological basis of the fishery for Caspian clupeids. Journal of Ichthyology, 26(2): 16-23.

Lalouie F., Nirani M., Taghavi M.J., Rezvani Gilkolaei S. (2006). The PCR-RFLP investigation of Clupeonella cultriventris from the south Caspian Sea. Iranian Journal of Fisheries Sciences, 15(2): 119-128. (In Farsi)

Lönnberg E. (1900). Short notes on Caspian fishes. Revue Internationale de Pêche et de Pisciculture, St. Petersbourg, 2(2): 3-6.

Mamedov E.V. (2006). The biology and abundance of kilka (Clupeonella spp.) along the coast of Azerbaijan, Caspian Sea. ICES Journal of Marine Science, 63(9): 1665-1673.

Mandych A.F. (1995). Enclosed seas and large lakes of eastern Europe and middle Asia. SPB Academic Publishing, Amsterdam. 273p.

Marammazi Gh. (1995). Survey of the biology of Tenualosa ilisha, southern Iran. Aquaculture Fishery Research Centre, Iranian Fisheries Research and Training Organization, Ahvaz. 212 p. (In Farsi)

Marammazi J., Al-Mukhtar M.A., Dehgan S., Maraashi S.Z., Eskandari G., Shakiba G., Mayiahi Y. (1995). Biology of soboor Tenualosa ilisha (phase 1), final report. Iranian Fisheries Research Organization, Tehran. 194p. (In Farsi)

Marammazi J., Al-Mukhtar M.A., Kiabi B.H. (1998). Comparison of morphometric and meristic traits and length frequency in soboor (Tenualosa ilisha Ham. Buch. 1822) of the migration pathways in Khuzestan province. Pajouhesh va Sazandegi, 11(39): 113-120. (In Farsi)

Marammazi J.G. (1994). Preliminary ecological investigation of six fish species in Zohre River. Iranian Fisheries Scientific Journal, 3(2): 57-70. (In Farsi)

Marammazi J.Gh. (1995). A study of identification and some biological aspects of sobour Tenualosa ilisha (Ham. Buch 1822) in Bahmanshir River, Khuzestan, Iran. M.Sc. Thesis, Tarbiat Modarres University. 78 p. (In Farsi)

Marammazi J.G.H., Kiabi B.O., Al-Mukhtar M.A. (1995). Morphometric and meristic characteristics of sobur Tenualosa ilisha (Ham. Buch 1822) in Bahmanshir River, Khuzestan, Iran. Iranian Fisheries Scientific Journal, 4(3): 40-55. (In Farsi)

Marammazi J.Gh., Al-Mukhtar M., Eskandari Gh.R. (1998). Fecundity estimation of sobour (Tenualosa ilisha, Ham. Buch. 1822) in Khouzestan Province's rivers. Iranian Fisheries Scientific Journal, 7(1): 69-82. (In Farsi).

Marammazi J.Gh., Al-Mukhtar M.A., Eskandari Gh.R., Dehghan S. (1996). Biological study of sobour Tenualosa ilisha (Ham. Buch. 1822) in migratory passways in southwest of Iran. Khuzestan Fisheries Research Centre, Ahaz, Iran. 214 p. (In Farsi)

Mazandarani M., Hajimoradloo A.M., Niazi E. (2016). Internal parasites of saposhnikovi shad, Alosa saposchnikowii (Grimm, 1887), from the southeastern part of the Caspian Sea, Iran. Iranian Journal of Fisheries Sciences, 15(3): 1067-1077.

Mikaili P., Shayegh J. (2011). An etymological review on fish common and scientific names in the Euphrates and Tigris. Research Journal of Fisheries and Hydrobiology, 6(4): 412-423.

Mikhailovsky A.A. (1941). On form-building in the Caspian herring Caspialosa brashnikovi (Borodin). Comptes Rendus (Doklady) de l'Académie des Sciences de l'URSS, nouvelle série, 30(6): 563-566.

Milton D.A., Chenery S.R. (2001). Can otolith chemistry detect the population structure of the shad hilsa Tenualosa ilisha? Comparison with the results of genetic and morphological studies. Marine Ecology Progress Series, 222: 239-251.

Moeeni S., Alizadeh Doughikolaie E., Mirvaghefi A.R., Moinie S. (2006). Soup powder production from kilka (Clupeonella engrauliformis) and a determination of its shelf life. Iranian Journal of Natural Resources, 59(1): 177-187.

Moeini S. (2002). Investigation on the method of production of sausages from kilka. Iranian Journal of Marine Science, 1(4): 111-120. (In Farsi)

Moeini S., Sabetian M., Khaleghi Gorj A., Farhangi M. (2009). An investigation on relationships of chemical indices of kilka (Clupeoenlla engrauliformis) with weight loss during cold storage at -18ºC. Iranian Scientific Fisheries Journal, 18(2): 129-140. (In Farsi).

Moghadamnia A.A., Mirhosseini N., Abadi M.H., Omranirad A.S., Omidvar S. (2010). Effect of Clupeenonella grimmi (anchovy/kilka) fish oil on dysmenorrhoea. Eastern Mediterranean Health Journal, 16(4): 408-413.

Mohamed A.R.M., Ali T.S., Hussain N.A. (2001). Stock assessment of hilsa shad Tenualosa ilisha in the Iraqi marine waters, northwest Arabian Gulf. Marina Mesopotamica, 16(1): 1-9.

Mohamed A-R.M., Qasim A.M.H. (2014a). Stock assessment and mangement of hilsa shad (Tenualosa ilisha) in Iraqi marine waters, northwest Arabian Gulf. International Journal of Fisheries and Aquaculture Studies, 1(52): 1-7.

Mohamed A-R.M., Qasim A.M.H. (2014b). A stock assessment and mangement of hilsa shad (Tenualosa ilisha) in Iraqi marine waters, northwest Arabian Gulf. World Journal of Fish and Marine Sciences, 6(2): 201-208.

Moini S., Koochekian A. (2003). Production of fish sauce from Caspian sea kilka, with use of traditional, microbial and enzymatic methods. Iranian Journal of Fisheries Sciences, 12(2): 79-94. (In Farsi)

Moradinasab A., Babaei N., Parsa M., Raeisi H., Daliri M., Rahnama B., Nekuro A., Vesaghi M.J. (2015). By-catch composition of kilka fishes (Clupeonella) lantern net in the southwest of the Caspian Sea (Bandar Anzali fishing grounds). The Third Iranian Conference of Ichthyology, Shiraz University, Shiraz, 6-7 May 2015. pp: 160.

Moradinasab G., Raeisi H., Ghorbani R., Paighamabari S. Y., Davoodi R. (2012). Weight-length relationship for Clupeonella cultriventris caspia (Svetovidov, 1941) in Iranian waters of the Southwest Caspian Sea. American Journal of Experimental Agriculture, 2(2): 240-246.

Motalebi A.A., Hasanzati Rostami A., Khanipour A.A., Soltani M. (2010). Impacts of whey protein edible coating on chemical and microbial factors of gutted kilka during frozen storage. Iranian Journal of Fisheries Sciences, 9(2): 255-264.

Motalebi A.A., Seyfzadeh M. (2011). Effects of whey protein edible coating on bacterial, chemical and sensory characteristics of frozen common kilka (Clupeonellia delitula (sic)). Iranian Journal of Fisheries Sciences, 11(1): 132-144.

Motamedzadegan A., Shahidi F., Mortazavi S.A., Pourazarang H., Hamzeh Sh., Shahidi Yasaghi S.A., Ghorbani Hasan Saraei A., Khanipour E. (2009). Effect of kilka fish myofibrillar protein hydrolysis by papain on peptide chain length and degree of hydrolysis. Journal of Agricultural Sciences and Natural Resources, 16(3): 172-181. (In Farsi)

Mousavi-Sabet H., Heidari A., Paknejad S. (2016). Length-weight and length-length relationships of the genus Alosa (Clupeoidei: Clupeiformes: Clupeidae) along the southern Caspian Sea coast. Journal of Applied Ichthyology, 32(1): 129-130.

Naderi M., Fazli H., Afraee M., Ganjian A. (1997). The study of reproduction, fecundity and diet in 3 species of kilka in the southern part of the Caspian Sea (Babolsar region). Iranian Fisheries Scientific Journal, 6(1): 65-78. (In Farsi)

Naem S., Mobedi I., Khomirani R., Abolghasemi S.J. (2002). A survey on gill parasites in cultured and wild fishes of western branch of Sepid-Rood River in Gilan province, Iran and introducing new species. Iranian Journal of Veterinary Research, Shiraz University, 3(2): 118-128. (In Farsi)

Najafpour N. (1997). Identification of some freshwater fishes of Khuzestan Province. Fisheries Research Centre of Khuzestan Province, Iranian Fisheries Research and Training Organisation, Tehran. 96 p. (In Farsi)

Najm M., Shokrzadeh M., Fakhar M., Sharif M., Hosseini S.M., Rahimi Esboei B., Habibi F. (2014). Concentration of heavy metals (Cd, Cr and Pb) in the tissues of Clupeonella cultriventris and Gasterosteus aculeatus from Babolsar coastal waters of Mazandaran Province, Caspian Sea. Journal of Mazandaran University of Medical Sciences, 24(113): 185-192.

Naseka A.M., Bogutskaya N.G. (2009). Fishes of the Caspian Sea: zoogeography and updated check-list. Zoosystematica Rossica, 18(2): 295-317.

Naseri M., Rezaei M., Abasi M. (2010). Lipid changes during long-term storage of canned common kilka (Clupeonella cultriventris). Journal of Fisheries (Iranian Journal of Natural Resources), 63(3): 229-241.

Naseri M., Rezaei M., Abasi M., Hosseini H., Jam S., Sabzevari O. (2009). Effects of pre-chilling process on the quality of muscle lipid and filling media in canned common kilka (Clupeonella cultriventris). Journal of Water and Soil Science (Journal of Science and Technology of Agriculture and Natural Resources), 12(46): 291-301.

Naseri M., Rezaei M., Hosseini H., Abbasi M., Sabzevari O. (2005). A comparison of conventional and fluorescence detection methods of cooking-induced damage to common kilka (Clupeonella cultriventris). Journal of Marine Sciences and Technology, 4(1-2): 75-82.

Naseri M., Rezaei M., Sabzevari O., Hosseini H., Mousapour M. (2006). Comparition impact of filling media on common kilka (Clupeonella cultriventris) canned quality by fluorescence detection. Iranian Journal of Food Science and Technology, 3(3): 37-48.

Nasri Tajan M. (2009). Determination of biological characteristics and blood groups of hilsa shad (Tenualosa ilisha) in the north of Persian Gulf (Bushehr coastal waters). Ph. D. Thesis, Islamic Azad University, Tehran. 128p. (In Farsi)

Nasri Tajan M., Keyvan A., Soltani M., Valinasab T., Vosoughi G.H. (2008). A survey of feeding behaviour of hilsa shad (Tenualosa ilisha) in the north of Persian Gulf (Bushehr coastal waters). Journal of Fisheries, 2(3): 43-50. (In Farsi)

Nassiri Moghadam H., Danesh Mesgaran M., Jahanian Najafabadi H., Jahanian Najafabadi R. (2007). Determination of chemical composition, mineral contents, and protein quality of Iranian kilka fish meal. International Journal of Poultry Science, 6(5): 354-361.

Navidshad B. (2014). Effects of fish (Clupeonella cultriventris, Caspian Sea originated) oil supplement on the serum lipoproteins and production of ǀwwǀ-3 fatty acids enriched broiler meat. Journal of Veterinary Research, 69(4): 405-414.

Nelson G., McCarthy L. (1995). Two new species of gizzard shads of the genus Nematalosa (Teleosti, Clupeidae, Dorosomatinae) from the Persian/Arabian Gulf. Japanese Journal of Ichthyology, 41(4): 379-383.

Nelson J.S., Grande T.C., Wilson M.V.H. (2016). Fishes of the World. Fifth Edition. John Wiley, Sons, Hoboken. 707p.

Nemati M., Javadian S.R., Ovissipour M., Keshavarz M. (2012). A study on the properties of Alosa (Alosa caspia) by-products protein hydrolysates using commercial enzymes. World Applied Sciences Journal, 18(7): 950-956.

Nikonorov I.V. (1964). Pump fishing with light and electric current, p. 577-579. In: Modern Fishing Gear of the World 2: Fishing News (Books), London. 603p.

Norouzi M., Nazemi A., Pourkazemi M. (2012). Population genetic study on common kilka (Clupeonella cultriventris Nordmann, 1840) in the Southwest Caspian Sea (Gilan Province, Iran) using microsatellite markers. African Journal of Biotechnology, 11(98): 16405-16411.

Norouzi M., Nazemi A., Pourkazemi M., Samiei M.H. (2013). Population genetic study on common kilka, Clupeonella cultriventris Nordmann, 1840, in the Southwest Caspian Sea, Gilan Province, using microsatellite markers. World Applied Sciences Journal, 21(11): 1675-1680.

Norouzi M., Nazemi A., Pourkazemi M., Samiei M.H. (2014). Analysis of genetic variability and differentiation of South Caspian Sea’s common kilka (Clupeonella cultriventris) using microsatellite markers in two different seasons. European Journal of Zoological Research, 3(1): 75-80.

Norouzi M., Nazemi A., Pourkazemi M., Samiei M.H., Ravai A. (2013). Population genetic structure of common kilka (Clupeonella cultriventris) in the south Caspian Sea coastline (Gilan Province) using microsatellite markers. Journal of Fisheries, 7(2): 69-76

Ojagh S.M., Sahari M.A., Rezaei M. (2004). Effects of natural antioxidants in quality of common kilka (Clupeonella cultriventris caspia) during storage with ice. Journal of Marine Sciences and Technology, 3(4): 1-7.

Osipov V.V., Kiyashko V.I. (2008). Method of age determination of Ponto-Caspian kilka Clupeonella cultriventris (Clupeiformes, Clupeoidei) using scales and otoliths. Journal of Ichthyology, 48(8): 637-643.

Ovissipour M., Rasco B., Shiroodi S.G., Modasnlow M., Gholami S., Nemati M. (2013). Antioxidant activity of protein hydrolysates from whole anchovy sprat (Clupeonella engrauliformis) prepared using endogenous enzymes and commercial proteases. Journal of the Science of Food and Agriculture, 93(7): 1718-1726.

Paknejad S., Heidari A., Mousavi-Sabet H. (2014). Morphological variation of shad fish Alosa brashnicowi (sic) (Teleostei, Clupeidae) populations along the southern Caspian Sea coasts, using a truss system. International Journal of Aquatic Biology, 2(6): 330-336.

Parafkandeh F. (2006). Effect of invasion of comb jelly Mnemiopsis leidyi on kilka fishery in the Caspian Sea. 6Th Iranian Congress of Marine Science and Technology, Tehran (abstract).

Parafkandeh F., Jamalzad F. (1997). A biological investigation into anchovy kilka (Clupeonella engrauliformis) in Anzali fishing ground. Iranian Fisheries Scientific Journal, 5(4): 31-42, 3. (In Farsi)

Parafkandeh Haghigi F. (1996). A look at kilka catch in 1995. Abzeeyan, Tehran, 24-27. (In Farsi)

Parafkandeh Haghigi F. (2010). Kilka population dynamics in the south of the Caspian Sea. Ph.D. Thesis, Islamic Azad University, Tehran. 128 p. (In Farsi)

Parafkandeh Haghighi F., Kaymaram F. (2010). Estimation of biological parameters of anchovy kilka Clupeonella engrauliformis (Borodin, 1904) in the South Caspian basin. Journal of Animal Environment, 2(3): 1-8. (In Farsi)

Parafkandeh Haghighi F., Kaymaram F. (2012). Significant changes in pelagic fish stocks of Clupeonella spp. in the south Caspian Sea. Iranian Journal of Fisheries Sciences, 11(3): 559-569.

Patimar R., Habibi S., Jafari F. (2011). A study on growth parameters of Alosa caspia caspia Eichwald, 1838 in the southern Caspian coast. Journal of Fisheries (Iranian Journal of Natural Resources), 64(1): 15-26.

Pavlov D.S., Reshetnikov Yu.S., Shatunovskiy M.I., Shilin N.I. (1985). Rare and disappearing fishes in the USSR and the principles of their inclusion in the "Red Book". Journal of Ichthyology, 25(1): 88-99.

Pillay S.R., Rosa H. (1963). Synopsis of biological data on hilsa Hilsa ilisha (Hamilton) 1822. Food and Agriculture Organization, Rome, Fisheries Biology Synopsis, 25: 61 p.

Poorgholam R., Sedov V., Yermalchev V., Besharat K., Fazli H. (1996). Stock assessment of kilka fishes by hydro-acoustic method 1994-95. Final Report, Mazandaran Fisheries Research Center, Sari. 125 p. (In Farsi)

Pourrafei L., Esmaeili H.R., Gholamhosseini A., Abbasi K., Valinassab T. (2016). Reconstructing phylogenetic relationships of Cluperiformes (sic) (Actinopterygii: Teleostei) based on the nuclear gene RAG1. The Fourth Iranian Conference of Ichthyology, Ferdowsi University of Mashhad, 20-21 July 2016, p. 42 (abstract).

Prikhod'ko B.I. (1979a). Relationship between abundance of young Caspian "anchovy kilka", Clupeonella engrauliformis, and water temperature. Journal of Ichthyology, 19(2): 86-95.

Prikhod'ko B.I. (1979b). Ecological features of the Caspian kilka (genus Clupeonella). Journal of Ichthyology, 19(5): 27-37.

Prikhod'ko B.I. (1981). Selective fishing for the Caspian "anchovy kilka", Clupeonella engrauliformis, using electric light. Journal of Ichthyology, 21(1): 104-110.

Pushbarnek E.B. (1987). Characteristics of the spawning population of Caspian shad, Alosa caspia caspia, from the Dagestan Coast during 1980-1984. Journal of Ichthyology, 27(5): 107-112.

Rahimi Bashar M.R., Alipour V. (2009). Survey of biometric characteristics and diet of anchovy kilka (Clupeonella engauliformis) (sic) in south-west of Capsian Sea. Journal of Biology Science, 3(2): 25-35. (In Farsi)

RaLonde R., Walczak P. (1971). A research proposal for studying the tyulka (Clupeonella sp.) populations of the southern Caspian Sea with notes on progress to date. Report of the Fisheries Research Institute, Bandar Pahlavi. MS, 6 p.

Razavi Sayad B. (1993). On kilka fishing status in the Caspian Sea. Iranian Fisheries Bulletin, 2: 11-25, 2-3. (In Farsi)

Reshetnikov Yu.S., Bogutskaya N.G., Vasil'eva E.D., Dorofeeva E.A., Naseka A.M., Popova O.A., Savvaitova K.A., Sideleva V.G., Sokolov L.I. (1997). An annotated check-list of the freshwater fishes of Russia. Journal of Ichthyology, 37(9): 687-736.

Rezaei M., Sahari M.A., Moeini S. (2007). Quality assessment of lipid in anchovy kilka (Clupeonella engrauliformis) during frozen storage at different temperature rates. Water and Soil Science (Journal of Science and Technology of Agriculture and Natural Resources), 10(4): 435-445. (In Farsi)

Rezaei M., Sahari M.A., Moini S., Safari M., Ghafari F. (2003). Comparative study on lipid quality on anchovy kilka (Clupeonella engrauliformis) under two temporary chilled transport storage methods. Iranian Journal of Fisheries Sciences, 12(3): 97-108. (In Farsi)

Rezaeian M., Sahari M.A., Moini S., Safari M., Rezaiean M., Ghafari F. (2002). Some qualitative characteristics of lipid in anchovy kilka, (Clupeonella engrauliformis), during frozen storage. Iranian Journal of Marine Sciences, 1(4): 55-64. (In Farsi)

Robins C.R., Bailey R.M., Bond C.E., Brooker J.R., Lachner E.A., Lea R.N., Scott W.B. (1991). World fishes important to North Americans exclusive of species from the continental waters of the United States and Canada. American Fisheries Society Special Publication, 21: 243 p.

Roohi A., Kideys A.E., Sajjadi A., Hashemian A., Pourgholam R., Fazli H., Ganjian Khanari A., Eker-Develi E. (2010). Changes in biodiversity of phytoplankton, zooplankton, fishes and macrobenthos in the southern Caspian Sea after the invasion of the ctenophore Mnemiopsis leidyi. Biological Invasions, 12(7): 2343-2361.

Roomiani L., Gholamhosein M., Khodadadi M. (2012). Estimation of growth parameters of hilsa shad, Tenualosa ilisha in Khuzestan Province using ELEFAN programme of FISAT II software. Journal of Marine Biology, 4(14): 11-20. (In Farsi)

Roomiani L., Jamili S. (2011). Population dynamics and stock assessment of hilsa shad, Tenualosa ilisha in Iran (Khuzestan Province). Journal of Fisheries and Aquatic Science, 6(2): 151-160.

Roushan Tabari M., Khoda Parast N.B., Vahedi F., Rostamian M.T. (2009). An investigation on the feeding behaviour of common kilka (Clupeonella cultriventris caspia) in the southern Caspian Sea, Mazandaran Province, Iran. Iranian Scientific Fisheries Journal, 18(3): 79-88. (In Farsi)

Sadeghi Bajgiran S., Pourkhabbaz A.R., Hasanpour M., Sinka Karimi M.H. (2016). A study on zinc, nickel, and vanadium in fish muscle of Alosa caspia and Sander lucioperca and food risk assessment of its consumption in the southeast of the Caspian Sea. Iranian Journal of Health and Environment, 8(4): 423-432. (In Farsi)

Sadough Niri A., Nikpour Y., Rajabzadeh E., Mahboobi Soofiani N.A., Ahmadi R. (2010). Assessment of heavy metals (Cd, Pb, Cu, Co, Ni) in some tissues of Tenualosa ilisha from northwest Persian Gulf and their relationship. Journal of Aquatic Sciences, 1(1): 73-87.

Sahari M.A., Ahmadnia A., Barzegar M., Noorolahi Z. (2014). Vitamin loss during frozen storage of Liza aurata (Risso, 1810), Cyprinus carpio L., 1758, Clupeonella cultriventris caspia (Nordmann, 1840), Rutilus frisii kutum (Kamenskii, 1901) and Sander lucioperca (L., 1758). Journal of Applied Ichthyology, 30(2): 366-371.

Salari A.M.A., Sadough N.A. (2009). Comparison of heavy metals content of muscle, liver and gill tissues of Tenualosa ilisha in Karoon River. The 6th Scientific Conference of Fisheries Resources, 3-4 March 2009, Basrah, pp: 136 (abstract).

Salmani A., Gholamipour S., Yousefian M. (2001). The changes in TVN and histamine of kilka within the different preservation methods. Iranian Journal of Fisheries Sciences, 10(2): 31-40. (In Farsi)

Salmani Joloudar A., Rezaeian M., Gholamipour S., Safari R., Pourgholam, R. (2009). Extraction of omega-3 fatty acid from kilka fish oil. Iranian Scientific Fisheries Journal, 17(4): 89-100. (In Farsi)

Sarhadi N., Motamedzadegan A., Taheri A., Azad M. (2012). Comparison study of the proximate composition and amino acid profile in the bones of goldstripe sardine (Sardinella gibbosa), anchovy kilka (Clupeonella engrauliformis) and Indian anchovy (Stolephorus indicus). Iranian Scientifc Fisheries Journal, 21(1): 101-112.

Sarhaddi N., Taheri A., Motamedzadegan A. (2015). Profiles in the bone of gold stripe sardine (Sardinella gibbosa), anchovy kilka (Clupeonella engrauliformis) and Indian anchovy (Stolephorus indicus). Journal of Fisheries (Iranian Journal of Natural Resources), 67(4): 535-545. (In Farsi)

Sayad Bourani M. S. (1999). Some biological survey of the Clupeonella engrauliformis (anchovy-like kilka) in Guilan coastal region, the south-west of Caspian Sea, in 1996-97. Iranian Journal of Fisheries Sciences, 8(1): 59-70. (In Farsi)

Sayyad Bourani M. (2003). Situation of kilka fishing in the Iranian part of the Caspian Sea. Iranian Inland Waters Aquaculture Center, Bandar Anzali. 15 p. (In Farsi)

Sayyad Bourani M., Abdolmalaki S., Khanipour A.A., Fazli H., Khedmati K. (2008). Catch composition and fishing trend of kilka in Iranian part of the Caspian Sea. Iranian Journal of Fisheries Sciences, 7(2): 73-86.

Sedov S.I., Rychagova T.L. (1983). Morphological characteristics of anchovy kilka, Clupeonella engrauliformis (Clupeidae), in winter and spring. Journal of Ichthyology, 23(3): 140-143.

Seyfzadeh M., Motalebi A.A., Kakoolaki S., Gholipour H. (2013). Chemical, microbiological and sensory evaluation of gutted kilka coated with whey protein based edible film incorporated with sodium alginate during frozen storage. Iranian Journal of Fisheries Sciences, 12(1): 140-153.

Shabanpour B., Moini S., Hamedi M., Poorkabireh M., Seyfabadi S.J. (2002). Effect of washing condition on organoleptic properties of surimi from kilka anchovy (Clupeonella engrauliformis). Iranian Journal of Marine Sciences, 1(3): 17-25. (In Farsi)

Shabanpour B., Shaabani A., Moeeni S., Poorkabireh M. (2006). The effect of different washing methods on chemical and gel forming properties of kilka surimi. Pajouhesh va Sazandegi, 19(3): 84-92. (In Farsi)

Shamsi S., Dalimi A. (1996). Identification of Pseudopentagramma symmetrica in Clupeonella sp. in the Caspian Sea. Pajouhesh va Sazandegi, 32: 104-105. (In Farsi)

Shamsi Sh., Dalimi A. (2001). Study on parasitic infection of Caspian common kilka. Proceedings of the 5th Symposium of Marine and Atmospheric Sciences, 7-9 March 2001. pp: 107. (In Farsi)

Shamsi Sh., Dalimi A., Jalali B., Alizadeh Shabani A. (1998). Incidence and intensity of helminths in Clupeonella grimmi Kessler 1877 (Clupeidae) from Caspian Sea. The 18th International Congress of Parasitology, 24-28 August 1998, Makuhari, Chiba, Japan, Parasitology International, 47(supplement): 170.

Shamsi Sh., Dalimi A., Pourgholam R. (1996a). Study on helminths of anchovy kilka. Proceedings of the 5th Iranian Biology Conference, University of Tabriz, Tabriz, Iran, 22-24 August 1996. (abstract)

Shamsi Sh., Dalimi A., Pourgholam R. (1996b). Helminths of Clupeonella engrauliformis from south coast of Caspian Sea, p. 50. In: Abstracts VII European Multicolloquium of Parasitology, 2-6 September 1996, Parma, Italy. Parassitologia, 38(1-2): 1-479.

Shamsi Sh., Dalimi A., Pourgholam R. (1997). Study on helminths of bigeye kilka, Clupeonella grimmi. Proceedings of the 6th Iranian Biology Conference, Shahid Bahonar University of Kerman, 25-27.pp: 67.

Shamsi Sh., Dalimi A., Pourgholam R. (1998a). Study on zoonotic parasites in kilka fish. Iranian Fisheries Scientific Journal, 7(1): 45-58. (In Farsi)

Shamsi Sh., Dalimi A., Pourgholam R. (1998b). Caspian kilka as a new host for some helminths. Proceedings of the 7th Iranian Biology Conference, University of Isfahan, Isfahan, Iran, 22-24 August 1998, pp: 138.

Shamsi Sh., Dalimi A., Pourgholam R., Habibi F. (1997). Parasitic study on Caspian kilka and its zoonoses. Proceedings of the 3rd Iranian Congress of Veterinary Students, University of Urmia, Urmia, Iran, 23-26 April 1997, pp: 83.

Sharma K.P. (1980). Further studies on the fish marketing conditions of Southern Iraq. The Arab Gulf, University of Basrah, 2(1): 223-228.

Shojaei A. (1998). Producing cracker from kilka. Iranian Fisheries Scientific Journal, 7(1): 25-44. (In Farsi)

Shojaei A.H. (1998). Cracker from kilka. Fishery Technology, Society of Fisheries Technologists (India), Kochi, 35(2): 90-94.

Shokrzadeh M., Ebadi A.G. (2005). Investigating and measurement of residues of D.D.T. and D.D.E. in four species of fishes in Caspian Sea, Iran. International Journal of Biology and Biotechnology, 2(1): 53-56.

Shokrzadeh M., Ebadi A.G. (2006). Investigating and measurement of residues of chlorobenzilate (organochlorine pesticides) in four species of the most consumed fishes in Caspian Sea (Iran). Pakistan Journal of Nutrition, 5(1): 68-70.

Shokrzadeh M., Saeedi Saravi S.S., Zehtab Yazdi Y. (2009). Lindane residues in cultivated cucumber and in the most consumed fish in Caspian Sea (Iran). Toxicology and Industrial Health, 25(8): 517-523.

Shokrzadeh Lamuki M., Saeedi Saravi S.S., Otadi N. (2012). Evaluation of residues of D.D.T. and D.D.A. in fish collected from Caspian Sea, Iran. Iranian Journal of Toxicology, 6(18): 704-708.

Shubina L.I. (1981). Growth and sexual maturation of the Caspian shad Alosa caspia caspia. Journal of Ichthyology, 21(2): 98-108.

Soleimani A., Varidi M.J., Sadeghi Mahoonak A.R., Nassiri Mahalati M. (2012). Study on chemical composition of Alosa caspia and evaluation of changes in its moisture and salt content during salting and drying. Iranian Food Science and Technology Research Journal, 8(2): 115-120.

Svetovidov A.N. (1945). On Clupeonella delicatula (Nordmann) of the Caspian and that of the Black and Azov seas. Comptes Rendus de l'Academie des Sciences de l'URSS, 46(5): 207-208.

Svetovidov A.N. (1952). Fauna of U.S.S.R. (Fauna SSSR). Fishes (Ryby). Vol. II, No. 1, Clupeidae (Sel'devye). Israel Program for Scientific Translations, Jerusalem (1963). 428p.

Svetovidov A.N. (1964). Systematics of the North American anadromous clupeoid fishes of the genera Alosa, Caspialosa, and Pomolobus. Copeia, 1964(1): 118-130.

Tavakoli H.R., Soltani M., Bahonar A. (2012). Isolation of some human pathogens from fresh and smoked had (Alosa kessleri) and silver carp (Hypophthalmichthys molitrix). Iranian Journal of Fisheries Sciences, 11(2): 424-429.

Tidwell J. (2001). Caspian calamities. Zoogoer, 30(3):2 p. (www.fonz.org/zoogoer/zg2001/30(3)caspianside. htm)

van den Eelaart A. (1954). Report to the Government of Iraq on the development of inland fisheries. Food and Agriculture Organization, Rome, EPTA Report, 270: 1-42.

Valipour Meri M., Alizadeh Doughikollaee E., Safari R., Arshadi A. (2011). Inhibitory effect of Bacillus licheniformis on G1 and G2 aflatoxins in culture media and kilka fish meal. Journal of Fisheries (Iranian Journal of Natural Resources), 64(2): 177-187.

Varshoie H., Mobedi I., Meshgi M.A., Jalali B. (2010). Survey on the digenean and monogenean helminthes of Clupeidae (Teleostes) (sic) from southern part of Caspian Sea. Research Journal of Parasitology, 5(3): 148-153.

Vetchanin V.I. (1984). Feeding of the Astrakhan shad, Alosa brashnikovi (Clupeidae), in the southeastern Caspian Sea. Journal of Ichthyology, 24(5): 143-147.

Vetchanin V.I. (1992). Razmer' i rost Brazhnikovskoi sel'di v yugo-vostochnom Kaspii [Size and growth of Brazhnikov's shad in the south-east Caspian]. Izvestiya Akademii Nauk Turkmenskoi SSR, Seriya Biologicheskikh Nauk, 1992(4): 48-53.

Vladykov V.D. (1964). Report to the government of Iran on the inland fisheries, especially of the Caspian Sea with special reference to sturgeon. Food and Agriculture Organization, Rome, Report FAO/EPTA 1818: 51 p.

Whitehead P.J.P. (1985). FAO species catalogue. Volume 7. Clupeid fishes of the world (suborder Clupeoidei). An annotated and illustrated catalogue of the herrings, sardines, pilchards, sprats, shads, anchovies and wolf-herrings. Part 1 - Chirocentridae, Clupeidae and Pristigasteridae. Food and Agriculture Organization, Rome, Fisheries Synopsis, 125, 7(1): 1-304.

Youssefi M.R., Hosseinifard S.M., Halajian A., Nasiri Amiri M., Shokrolahi S. (2011). Pronoprymna ventricosa (Digenea: Faustulidae) in Alosa caspia fish in north of Iran. World Journal of Fish and Marine Sciences, 3(2): 104-106.

Zahmatkesh M., Shabanipour N., Zahmatkesh A., Abbasi K. (2014). Seasonal changes in diet of braschnikowi shad (Alosa braschnikowi) from eastern coast of Guilan Province. Second National Congress of Iran Fisheries and Aquatic Sciences, Islamic Azad University of Bandar Abbas (abstract). (In Farsi)

Zahmatkesh M., Shabanipour N., Zahmatkesh A., Abbasi K. (2016). Population structure of Alosa Linck, 1970 (sic) in the southern coast of Caspian Sea (Guilan Province). Journal of Animal Researches, 28(4): 457-465. (In Farsi)

Zakharyan G.B., Teruni V.N. (1979). The influence of light-assisted fishing of the "kilka" of the genus Clupeonella on stocks of the Caspian shad of the genus Alosa (Family Clupeidae). Journal of Ichthyology, 19(5): 129-132.

Zamakhaev D.F. (1944). K voprosu o sistematicheskom polozhenii prokhodnykh sel'dei Kaspiya [On questions concerning the systematic position of the Caspian anadromous herrings]. Zoologicheskii Zhurnal, 23(2-3): 65-81.

Zamani A., Rezaee M., Madani R. (2012). In-vitro effects of biochemical factors on trypsin activity from intestine and pyloric caeca of common kilka (Clupeonella cultriventris caspia) for inhibition of belly bursting. Iranian Scientific Fisheries Journal, 20(4): 53-62. (In Farsi)

Zare P., Kasatkina S.M., Shibaev S.V., Fazli H. (2017). In situ acoustic target strength of anchovy kilka (Clupeonella engrauliformis) in the Caspian Sea (Iran). Fisheries Research, 186(1): 311-318.

Refbacks

  • There are currently no refbacks.